|“||I lost my family||„|
|~ Speckles explaining his actions|
Speckles was part of the specialized all-animal team of hopeful FBI agents-to-be, helping to infiltrate Leonard Saber's mansion to retrieve important data on a weapon codename known as Clusterstorm. During the climax of the film, Speckles revealed to Darwin that he—using the pseudonym of "Yǎnshǔ", Chinese for "Mole"—was the one who created Clusterstorm by manipulating Leonard Saber and that his family and home had been destroyed by humans. As a result, Speckles vowed to get revenge on humans by bombarding the Earth with space junk and debris to drive them underground.
At the near end of the film, Darwin manages to break Speckles free from his insanity by saying that even though his family is gone, Speckles is still a part of the G-Force team, which had become his family, one he was close to destroying like what had happened to his old family long ago. Horrified by this realization, Speckles tries to shut down the Clusterstorm initiative and the robot he created but fails to do so. Luckily, Darwin manages to destroy the robot by inserting a virus-infected PDA. He is also saddened when the team finds Hurley and believe him dead, saying that it's all his fault for an innocent life to have died under his blind obsession for revenge, and is later relieved when they learn that Hurley had survived.
At the end of the film, Speckles is assigned into removing the malicious chips inside thousands of Saberling appliances but promises to join the team again once he was done if they would have him.