Lost children of my children's children, bickering, whining, endlessly skirmishing metal playthings, behold, for the fire has arrived. You have enjoyed a millennia of recess in the absence of your pedagogues. You have spread your banal sparks across the blackness of space and now, your curfew is nigh. God walks among you now. I am Megatronus.
~ Megatronus revealing himself to the Transformers.
Megatronus Prime is the ultimate antagonist of Transformers: Titans Return. He is a member of the original Thirteen Primes of Cybertron, as well as Megatron's namesake.
He is voiced by Mark Hamill, who is also known for portraying the JokerinBatman: The Animated Seriesand the Batman Arkham games.
After Starscream was defeated by the combined forces of Windblage, Metroplex, Optimus and Megatron, Megatronus found his spark and placed it in the body of Trypticon.
After Trypticon was defeated by the power of the Matrix of Leadership and the Enigma of Combination was freed, Megatronus appeared and claimed the two artifacts. Megatron attempted to stall and distract Megatronus by praising him while Optimus and Fortress Maximus sneak attacked him, but Megatronus avoided both attacks, killing Optimus and tearing off Maximus's arm.
With the Matrix of Leadership and Enigma of Combination in his grasp, Megatronus bid farewell to the Transformers before teleporting away.