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In general, they have their own motivation but they all have in common a mission to bring Clementine to Malmoth.
When they fail, usually dying, Malmoth punishes them by transforming them into crawling creatures.
They keep their human head, but end up with an animal body, often reptile.
A conversation between Count Cafardo and Malmoth suggests that they had this form before their master sends them to do evil on Earth.