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Jacob Lane is a Lycan (werewolf) hybrid creature and the main antagonist in Underworld: Awakening. He is the director of a biotech pharmaceutical company called Antigen. He is portrayed by Stephen Rea, who previously played the character of the Groom (also a werewolf) in the film The Company of the Wolves.
His wife is also mentioned as having chosen to remain human. Lane considers this as her 'abandonment' of himself and their son. However, this theory is only suggested, rather than being stated. It is also possible that he and Quint were accidentally infected with Lycanthropy, and then attempted to force Lane's wife into becoming a Lycan as well. He is later revealed to be part of the Lycans' master plan to eradicate their Vampire enemies.
While Quint is fighting Selene, Lane turns to his Hybrid form and almost kills Sebastian when he tries to prevent him from kidnapping Eve, but before he can do so. They appear evenly matched until Eve grabs Lane and slams him into the windshield of a car. She then rips out his jugular vein, killing him slowly.