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Appearance and Abilities
The Ape Titan is 17m in stature, larger than the common 15m Titan. While all other Titans seen in the story thus far have been distinctly human in appearance, the Ape Titan possesses more animal-like features. It has apelike features, and most of its body except for the face and torso is covered in fur.
The Ape Titan possesses numerous abnormal traits. Most notable are its high intelligence and its ability to speak human language. Possibly as a consequence of this, it is capable of controlling other Titans through a combination of verbal orders and physical intimidation. Additionally, the Ape Titan possesses the strength to throw heavy projectiles great distances with excellent accuracy and the dexterity to scale the Wall Maria.
The Beast Titan is a Titan Shifter. In his human form he is well toned, has wavy blonde hair along with a beard and mustache. He also wears glasses.
He is revealed to possess some unknown means to transform humans into Titans. These Titans are able to function at night, unlike regular Titans.
Clash of the Titans arc
The Beast Titan makes his first appearance when Titans are mysteriously appearing within Wall Rose even though the Wall remains intact. He is encountered by Mike Zacharius, who instantly notes the strangeness of his characteristics, mistaking the Titan with a regular Deviant-type. The Titan displays an unexpected behavior when he grabs Mike's horse and launches it at him, preventing his escape and throwing him from the roof he was on.
After Mike falls to the mouth of another Titan, the Beast Titan displays the ability to speak, ordering the other Titans to not eat him so that he can question Mike about the Three Dimensional Maneuver Gear. However, the Titan disobeys him and the Beast Titan crushes its head to release Mike.
The Beast Titan then proceeds to interrogate Mike, but as he neglects to respond out of shock, he theorizes that Mike does not speak due to them not speaking the same language, or Mike being too afraid. After inspecting the gear, he allows the surrounding Titans to devour Mike. As Mike screams in horror, the Beast Titan is amused to see that they did actually speak the same language after all. He finally leaves the scene contemplating the 3D Maneuver Gear.
He next appears walking past Castle Utgard, seemingly ignoring those taking refuge within. Instead, he uses his incredible strength to scale Wall Rose and then disappears.As thesoldiers manage to fend off an attack, he suddenly turns his attention towards them. He begins to break off chunks of the Wall, hurling them long distances at the members of the Survey Corps trapped in Castle Utgard. The first shot kills their horses, and the second kills Lynne and Henning.He then howls, summoning more Titans to continue the assault on the castle. Nanaba angrily notes that he seems to be intentionally toying with them. Afterwards, he climbs over the Wall and disappears into the lost territory of Wall Maria.
When Ymir and Eren Yeager are kidnapped by Reiner Braun and Bertolt Hoover, Ymir states that the Beast Titan is directly responsible for the appearance of Titans within Wall Rose; possibly in order to "test" humanity's strength. She also notes that the reaction of Reiner and Bertolt to his appearance resembled gawking, excited children. She theorizes that locating him must be one of their primary goals and that it is key for them to return to their village.
The Uprising arc
In the aftermath of Eren's failed kidnapping, an investigation is performed at the village of Ragako. The investigation confirms the suspicion that the villagers had been transformed into Titans, though the methods employed by the Beast Titan remains undisclosed.
Two months after Historia Reiss is crowned queen, the Beast Titan is shown in Shiganshina District defeating the Armored Titan in battle. While Bertolt takes out Reiner's body from his Titan form, the human within the Beast Titan emerges from his Titan body, telling Bertolt and Reiner to worry about Annie's rescue later. He then reveals his plan to ambush the Survey Corps in Shiganshina with their help in order to capture Eren and steal the Coordinate, their top priority.
Return to Shiganshina arc
After some time, the Survey Corps initiate the operation to retake Wall Maria. After Eren manages to seal the outer gate of Shiganshina, the Beast Titan, Reiner and Bertolt launch their operation. After Reiner's failed attack and its momentary downfall, the Beast Titan transform along with a numerous horde of Titans outside the Wall. After that, he picks up a giant boulder and launches it at the inner gate, so that horses could not get over the hole, keeping the soldier from running away.
The Beast Titan then roars, ordering small Titans to attack Survey Corps horses, while larger type of Titans form a circle around the entrance of Shiganshina district in order to prevent anyone from escaping the district.
- The Beast Titan's appearance resembles several ape-like cryptids, such as the Sasquatch/Bigfoot, Yeti, and Yowie.
- The Beast Titan is the first Titan fully capable of advanced communication, speaking the human language effortlessly. It also shows greater dexterity and fine-motor control, allowing it to remove Mike Zacharius's gear without damaging it.
- It appears to have had limited or no contact with humans inside the Walls for at least half a century, as it expresses interest in humanity's Three Dimensional Maneuver Gear. It also wonders whether or not it speaks the same language as Mike, though it's unknown whether this was genuine or simply mocking his terrified state.